-DPhil at Oxford University (1999-2003), Supervised by Prof. Louise Johnson
-Post-doctoral work with Elena Conti initially at EMBL Heidelberg (2004-2007) and later at the MPI for biochemistry in Martinsried (2007-2010)
-MRC career development Fellow at Wellcome Trust centre for Cell Biology in Edinburgh (2011-2016)
-WT Senior Research Fellow at Wellcome Trust centre for Cell Biology in Edinburgh (2016-present)